While raiding Berk’s Squab Farm, Mr. Fox triggers a fox trap, caging himself along with his wife, Felicity. She reveals her pregnancy to her husband and pleads with him to find a safer job if they escape, and he agrees.
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Two years (12 fox-years) later, the Foxes and their son Ash are living in a hole. Mr. Fox, now a newspaper columnist, moves the family into a better home inside a tree, ignoring the warnings of his lawyer Clive Badger about how dangerous the area is for foxes due to its proximity to facilities run by three farmers: Walt Boggis, Nate Bunce, and Frank Bean.
Angered by the raids, the farmers decide to kill Mr. Fox. They camp out near his home, waiting for the Foxes to surface for food and water. Underground, Felicity is upset that Mr. Fox has returned to his thieving ways.
Mr. Fox organizes a rescue mission to save Kristofferson held captive by the farmers. With the help of the local animal residents, they launch a counterattack, allowing Mr. Fox, Ash, and Kylie to enter Bean Annex undetected.
As the animals settle into their new homes in the sewers, Mr. Fox raids a supermarket owned by the farmers, revealing Felicity’s upcoming pregnancy. The animals celebrate their abundant new food source.
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