During the summer, the Heffleys plan a road trip to Meemaw’s 90th birthday party. Greg and Rodrick are not thrilled about the idea, but Susan promises to make it fun for everyone. Their journey begins with a stop at a restaurant called Corny’s.
Greg ends up in a ball pit with a diaper stuck to his hand while attempting to rescue Manny. The incident goes viral online, and Greg becomes known as “Diaper Hands”. Determined to leave the embarrassing meme behind, he hatches a plan to attend the Player Expo gaming convention.
Despite Susan’s ban on technology during the road trip, Greg manages to sneak his phone and reveals his plan to attend the Player Expo to Rodrick. They decide to go together, with Rodrick hoping to win money for his band’s van.
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While on the road, the Heffleys encounter the Beardos, a family with whom Greg gets into a conflict. Mr. Beardo seeks revenge on Greg, leading to a series of misadventures during the trip.
The Heffleys attend a country fair where Manny wins a piglet, leading to chaos in the car. Greg redirects their GPS to the Player Expo, sparking a search for the missing boys.
Greg and Rodrick attend the Expo, but things take a turn when Susan arrives and reveals Greg’s embarrassing past to his favorite internet star. A confrontation ensues, but ultimately, the family learns to reconcile and work together.
As the journey concludes, the Heffleys face challenges such as a stolen scrapbook and a pursuit by the Beardos. Through a series of events, they manage to retrieve their belongings and arrive at Meemaw’s party in time.
Despite the ups and downs of the trip, the family bonds and looks forward to future adventures. Greg embraces his “Diaper Hands” fame, finding happiness in unexpected moments.